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Candida rugosa lipase N-acetylglucosamine 2-epimerase NMR spectroscopy Neu5Ac analogues PEG Q1-390 QD1-999 QD450-801 Thiamin diphosphate X-ray crystallography alkaline pH artificial metalloenzyme asymmetric hydrogenation asymmetric synthesis baker’s yeast biocatalysis chemoenzymatic chemoenzymatic synthesis chloroketones classification of OYE cofactor analogues covalent immobilization d-lysine decarboxylase deprotonation/reprotonation mechanism dynamic kinetic resolution enantioselective bioreduction enzyme evolution enzyme stabilization halohydrins